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Each spinal and cranial motor nerve cell, through the extensive arborization of the terminal part of its efferent ber, comes into contact with only a few or up to 1000 or more muscle bers; together, the nerve cell, its axons, and the muscle bers they subserve constitute the motor unit All variations in the force, range, rate, and type of movement are determined by the number and size of motor units called into action and the frequency and sequence of ring of each motor unit Feeble movements involve relatively few small motor units; powerful movements recruit many more units of increasing size Within a few days after interruption of a motor nerve, the individual denervated muscle bers begin to contract spontaneously This isolated activity of individual muscle bers is called brillation Inability of the isolated ber to maintain a stable membrane potential is the likely explanation Fibrillation is so ne that it cannot be seen through the intact skin, but it can be recorded as a small, repetitive, short-duration potential in the electromyogram (EMG) (Chap 45) When a motor neuron becomes diseased, as in progressive spinal muscular atrophy, it may manifest increased irritability, ie, the axon is unstable and capable of ectopic impulse generation, and all the muscle bers that it controls may discharge sporadically, in isolation from other units The result of contraction of one or several such motor units is a visible twitch, or fasciculation, that appears in the EMG as a large spontaneous muscle action potential Simultaneous or sequential spontaneous contractions of multiple motor units cause a rippling of muscle, a condition known as myokymia If the motor neuron is destroyed, all the muscle bers that it innervates undergo profound atrophy termed denervation atrophy The motor nerve bers of each ventral root intermingle with those of neighboring roots to form plexuses, and although the muscles are innervated roughly according to segments of the spinal cord, each large muscle comes to be supplied by two or more roots In contrast, a single peripheral nerve usually provides the complete 39.

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the short option premium may have the potential to be reduced against the long options during the term of the option spread To demonstrate the iron butter y, let s look at the SPDR S&P 500 exchange-traded fund (SPY) option chain shown in Figure 85 In the gure we can see that the SPY is trading at 8426 which makes the 85 call and put the at-the-money options If we consider selling the May SPY 85 call option for 400 and the May SPY 85 put option for 470, we capture a total short premium of 870 Along with the short we would purchase out-of-the-money cover positions for each option by buying the May SPY 95 call option for 72 and the May SPY 75 put option for 146, thus giving us a total premium capture of 652 The combined theta on the trade is the key to success with the iron butter y You should consider the iron butter y for short-term expiration options to take advantage of the accelerated time value under the 60-day mark In this case the combined theta of the shorts is greater than 100, while the combined theta being lost daily is around 70 The iron butter y takes advantage of time value decay It also takes advantage of decreases in vega during the term of the trade If implied volatility collapses, the short option

motor innervation of a muscle or group of muscles For this reason, paralysis due to disease of the anterior horn cells or anterior roots has a different pattern than paralysis following interruption of a peripheral nerve These patterns follow the distribution shown in Table 46-1, page 1118 For example, section of the L5 motor root causes paralysis of the extensors of the foot and a foot drop, while a lesion of the peroneal nerve does the same but does not affect the invertors of the foot that are also innervated by L5 but via the tibial nerve All motor activity, even the most elementary re ex type, requires the synchronous activity of many muscles Analysis of a relatively simple movement, such as clenching the st, conveys some idea of the complexity of the underlying neuromuscular arrangements In this act the primary movement is a contraction of the exor muscles of the ngers, the exor digitorum sublimis and profundus, the exor pollicis longus and brevis, and the abductor pollicis brevis In the terminology of Beevor, these muscles act as agonists, or prime movers For exion to be smooth and forceful, the extensor muscles (antagonists) must relax at the same rate as the exors contract (reciprocal innervation, or Sherrington s law) The muscles that ex the ngers also ex the wrist; since it is desired that only the ngers ex, the extensors of the wrist must be brought into play to prevent its exion; they are synergists Lastly, during this action of the hand, appropriate exor and extensor muscles stabilize the wrist, elbow, and shoulder; muscles that accomplish this serve as xators The coordination of agonists, antagonists, synergists, and xators is effected mainly by segmental spinal re exes under the guidance of proprioceptive sensory stimuli In general, the more delicate the movement, the more precise must be the coordination between agonist and antagonist muscles All voluntary ballistic (phasic) movements to a target are effected by the activation of ensembles of motor neurons, large ones supplying large motor units and small ones, small motor units The smaller ones are more ef ciently activated by sensory afferents from muscle spindles, more tonically active, and more readily recruited in re ex activities, postural maintenance, walking, and running The large motor units participate mainly in phasic movements, which are characterized by an initial burst of activity in the agonist muscles, then a burst in the antagonists, followed by a third burst in the agonists The strength of the initial agonist burst determines the speed and distance of the movement, but there is always the same triphasic pattern of agonist, antagonist, and agonist activity (Hallett et al) The basal ganglia and cerebellum set the pattern and timing of the muscle action in any projected motor performance These points are discussed further in Chaps 4 and 5 Unlike the phasic movements just described, certain basic motor activities do not involve reciprocal innervation In support of the body in an upright posture, when the legs must act as rigid pillars, and in shivering, agonists and antagonists contract simultaneously Locomotion requires that the extensor pattern of re ex standing be inhibited and that the coordinated pattern of alternating stepping movements be substituted; the latter is accomplished by multisegmental spinal and brainstem re exes, the so-called locomotor centers Suprasegmental control of the axial and proximal.

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